中文摘要：

Marattioid 蕨纲植物是首先出现在中间的原始脉管的植物的一个古老的系含碳。现存成员几乎只被限制为热带区域，并且充满种类的家庭 Angiopteridaceae 在他们到东方半球的分发是有限的；在这个组以内的关系当前是含糊的。这里在 Angiopteris Hoffm 之间的种系发生的关系。并且 Archangiopteris 基督 et Gies。被评估基于叶绿体 rbcL 基因和 trnL-F 的在顺序分析内部有 MEGA2 和 MrBayesv3.0b4 的遗传因子的分隔符。根据种系发生的模式和石块记录，我们进一步为二个类估计了分叉时间。种系发生的树表明在这研究的 Angiopterisand Archangiopteris 的所有种类与强壮的统计支持形成了一个单音的门的组，但是在二个类之间的关系基于单个顺序分析仍然保持未解决。在另一方面，联合数据的顺序分析设定第一表明那 Archangiopteris 种分叉，当传统地假定，显示 Archangiopteris 不能是直接衍生物。Angiopteris 和 Archangiopteris 的 clade 在迟了的渐新世显得有多样化(≈ 2 6 妈) 基于分子的估计。因此， extantAngiopteris 和 Archangiopteris 的进化历史被古老的起源和最近的多样化描绘了，并且这些组不是遗物并且当传统地考虑了，危及系。

英文摘要：

Marattioid ferns are an ancient lineage of primitive vascular plants that first appeared in the middle Carboniferous. Extant members are almost exclusively restricted to tropical regions, and the specles-rich family Angiopteridaceae are limited in their distribution to the eastern hemisphere; relationships within the group are currently vague. Here the phylogenetic relationship between Angiopterls Hoffm. and Archangiopteris Christ et Gies. was evaluated based on the sequence analysis of chloroplast rbcL gene and trnL-F intergenic spacer with MEGA2 and MrBayes v3.064. On the basis of the phyIogenetic pattam and fossil record, we further estimated the divergence time for the two genera. The phylogenetic trees revealed that all species of Angiopteris and Archangiopteris in this study formed a monophyletic group with strong statistical support, but the relationship between the two genera remined unresolved based on individual sequence analysis. On the other hand, the sequence analyses of combined data set revealed that Archangiopteris species diverged first, indicating that Archangiop teris may not be a direct derivative as traditionally assumed. The clade of Angiopterls and Archangiopteris appears to have diversified in the late Oligocene （=26 Ma） based on the molecular estimate. Thus, the evolutionary history of extant Angiopteris and Archangiopteris has been characterized by ancient origin and recent diversification, and these groups are not relic and endangered lineages as traditionally considered.