中文摘要：

Biographical note of the first author： Hao ZHANG， male, born in 1981, Ph. D candidate, majoring in cell biology and proteomics Correspondence： Dr. Da-Cheng HE, Beijing Normal University, Key Laboratory of Cell Proliferation and Regulation of Ministry of Education, 19th Xinjiekouwai St. 100875, Beijing China. Fax： 8610-58805042. E-mail： dhe@bnu.edu.cn Correspondence： Dr. Xue-Yuan XIAO , Beijing Normal University, Key Laboratory of Cell Proliferation and Regulation of Ministry of Education, 19th Xinjiekouwai St. 100875, Beijing China. Fax： 8610-58805042. E-mail： xyxiao@bnu.edu.cn