晚石炭世莫斯科期刺毛类(Chaetetids)是重要的造礁生物,在全球范围内广泛分布并建造礁体。中国已报道的刺毛类礁主要分布在辽东太子河流域,产在上石炭统本溪组海相灰岩中。刺毛类生长形态多样,有层状、板状、丘状、分枝状和柱状,主要以板状覆盖沉积物的方式造礁。研究区刺毛类礁中刺毛类生长形态及其所主导的生物群落在垂向上变化明显,由此将礁体的建造划分为三个阶段:①定殖阶段:刺毛类较少,生长方式单一,多以层状形态生长在棘屑滩上。层状刺毛类体积小,抗浪能力较弱,很难在其周围创造水流平缓的可供其他底栖生物生存的环境。因而该阶段群落生物种类单一,数量稀少;②繁盛阶段:刺毛类丰富,生长形态多样,以板状、分枝状和丘状为主。板状刺毛类单独形成的礁骨架,或与分枝状和丘状刺毛类组合起来的礁骨架,都能在其周围一定范围营造适宜底栖生物生存的环境,因此该阶段群落的生物多样性高;③衰退阶段:刺毛类很少,多为薄层状,断续生长在棘屑组成的基底上。薄层状刺毛类很脆弱,基本失去改善周围环境的能力,因此该阶段群落生物多样性很低。以上分析说明刺毛类生长形态的变化可以影响生物群落的组成和演化。生物群落的演化过程就是礁体建造的过程。从刺毛类生长形态的角度研究礁体建造过程,为分析由具有多种生长形态的造礁生物建造的生物礁的成礁机制提供新思路。
Chaetetids, an important reef-building organism, were globally distributed in the Late Carboniferous (Moscovian). Chaetetids reefs reported in China develop mainly in marine limestone of the Benxi Formation (Upper Carboniferous) in the Taizi River Basin, East Liaoning. Chaetetids have a variety of growth forms, such as laminar, tabular, moundy, branched and columnar. Chaetetids reefs were built in the form of tabular cover. In the study area, the reef-building process can be divided into three stages, according to the obvious changes of Chaetetids growth forms and the community controlled by Chaetetids in the vertical direction. (1)Colonization stage. There were a few Chaetetids which mainly were laminar colonizing the crinoids debris bank. It's difficult to create the stable environment for other benthic organisms living, because laminar Chaetetids were small and had a weak ability to resist wave. Hence, the community was simple in the stage. (2)Flourish stage: Chaetetids were abundant and their growth forms included tabular, columnar, moundy and branched. The reef framework built by tabular Chaetetids or the other forms could create the stable environment for other benthic organisms living. Therefore, the diversity of the community was high in the stage. (3) Decline stage: There were few Chaetetids. They occurred as discontinuous-laminar forms colonizing the substrate which composed of crinoids debris. Laminar Chaetetids were fragile and lost ability to improve the environment. In the stage, the community was simple. Overall, Chaetetids growth forms can influence the formation and succession of the community. Chaetetids reef was built as the community evolved. The study focuses on Chaetetids growth forms and reef-building process. It can provide a new perspective to understand the reef-building mechanism.