记述了河南确山后胥山晚上新世裂隙堆积物中新发现的鲁南鼩鼱Lunanosorex材料;重新观察了内蒙古比例克的李氏鲁南鼩鼱相似种Lunanosorexcf.Llii标本。确山鲁南鼩鼱的形态特征与属型种L.lii较为一致,其较丰富的材料为了解该种的形态变异及特征补充提供了新的依据。内蒙古比例克的鲁南鼩鼱与属型种有较大的差异,被确定为新种L.qiui sp.nov.。根据比例克新种和确山新材料的研究,修订了Lunanosorex的属征和Beremendiini族征的定义,探讨了鲁南鼩鼱与东亚地区发现的其他牙齿染色的大型鼩鼱类之间的系统关系,阐证了日本中更新世的Shikamainosorex与北美上新世的Paracryptotis关系不大,不应归人Blarrinini族,而与亚洲大陆的鲁南鼩鼱有较近的亲缘关系,应当属于Beremendiini族。
Lunanosorex is one of important insectivore members of Neogene micromammal fauna in North China. It was established by Jin and Kawamura in 1996, based on only one mandible from the Late Pliocene fissure deposit in Qipanshan Hill, Yinan County, Shandong Province. Thereafter, it was reported in Bilike and Gaotege, Nei Mongol ( Qiu and Storch, 2000; Li et al., 2003 ). In 2004, more Lunanosorex specimens were excavated by a team of the IVPP from Queshan County, Henan Province. On the basis of the evolutionary stage of micromammals, the fossil bearing deposits of the Queshan County is possibly the Late Pliocene in age. With a detailed study of the aforementioned specimens, the revision of the genus and species and the relationship among some of the large-sized shrews are discussed in the present paper. 1 Systematic paleontology Order Insectivora Bowdich, 1821 Family Soricidae Gray, 1821 Subfamily Soricinae Fischer von Waldheim. 1817 Tribe Beremendiini Reumer, 1984 Genus Lunanosorex Jin & Kawamura, 1996 Type species Lunanosorex lii Jin & Kawamura, 1996. Emended diagnosis Medium- to large-sized soricid. Coronoid process long and broad, external temporal fossa deep and coronoid process spicule well developed; interarticular area broad, lower facet of condyle shifted anteriorly; internal temporal fossa deeply pocketed and continuing to mandibular foramen through a medium-sized opening; the upper pterygoid fossa shallow; teeth pigmented; upper incisor unfissident; P4 with developed protocone; upper P4 and molars with narrow talon; lower incisor with tricuspulate or bicuspulate; p4 with narrow and deep posterolingual basin; ml and m2 with developed entoconid crest; and m3 reduced. Lunanosorex lii Jin & Kawamura, 1996 ( Fig. 2) Holotype IVPP V 10813, a left mandible. Referred specimens IVPP V 14973.1 -2, two nearly completed right mandibles with i and p4 - m3 ; V 14973.3 - 4, two nearly completed mandibles with p4 - m3 and ml - m3, respectively; and V 14973.5, a right mandible with broken