光合作用对光和CO2响应模型是研究植物生理和植物生态学的重要工具,可为植物光合特性对主要环境因子的响应提供科学依据。该文综述了当前光合作用对光和CO2响应模型的研究进展和存在的问题,并在此基础上探讨了这些模型的可能发展趋势。光合作用涉及光能的吸收、能量转换、电子传递、ATP合成、CO2固定等一系列复杂的物理和化学反应过程。光合作用由原初反应、同化力形成和碳同化3个基本过程构成,任一个过程均可对光合作用速率产生直接的影响。光合作用对光响应模型只涉及光能的转换,而光合作用的生化模型包含了同化力形成和碳同化这两个基本过程。把光合作用的原初反应,即把参与光能吸收、传递和转换的捕光色素分子的物理参数(如捕光色素分子数、捕光色素分子光能吸收截面、捕光色素分子处于激发态的平均寿命等)结合到生化模型中,可能是今后光合作用对光响应机理模型的发展方向。
The light and CO2 response curve of photosynthesis is an important tool to study plant physiology and plant ecology that can provide a scientific basis for the response of plant photosynthetic properties to environmental factors. This review considered the progress and potential weaknesses of light and CO2 response models of photosynthesis and discussed research trends. Photosynthesis,which involves energy of light,absorption,energy conversion,electron transfer,ATP synthesis,CO2 fixation etc.,is a complex physical and chemical reaction process. It includes three basic steps:the primary reaction,the assimilatory power forms and the carbon assimilation,and each link may directly influence other processes. Classical models on photosynthetic light response only involve with light energy absorption,and biochemistry models do with the assimilatory power to form as well as carbon assimilation. A future direction of research of the mechanistic model of photosynthetic light response is the primary reaction of photosynthesis,namely participation the energy of light absorption,the transmission and the transformation of the harvesting light pigment member the physical parameter (e.g.,the light-harvesting pigment mole-cules,light energy absorption cross-section of the harvesting pigment,the mean lifetime of the harvesting light pigment) unify in the biochemistry model.