华南地区中泥盆世晚期发生了一次大海侵,而块状群体的内板珊瑚(Endophyllum)则是这次大海侵开始时的标志性化石,Endophyllum往往与其它的群体造礁珊瑚(如Argutastrea,Alveolites等)和层孔虫(如Strom-atopora,Clathrostroma等)形成生物层或生物岩礁。华南地区Endophyllum的地层层位十分稳定,大致位于牙形类的Polygnathus varcus带的下部,它经常与吉维特期标准腕足类鸮头贝(Stringocephalus)共生。根据对产自新疆北部东准噶尔和西准噶尔所谓的"Endophyllum"的外部形态特征和内部构造的微细结构的深入研究,并将它与华南地区湘、桂、黔、滇、川等省(区)所产出的Endophyllum进行仔细的对比之后认为:两者存在着很大的差别,它们应分属于不同的属、甚至不同的科或科以上的分类单元,而且它们产出的地层层位也不尽相同,北疆所谓的"Endophyllum"要比华南的Endophyllum出现得早。
The South China Plate was formed by collision of the Yangtze and Cathaysian plates as a result of the Caledonian Movement.At the beginning of the Early Devonian,most of South China was above sea level,apart from the deep Qinzhou trough and Qujing remnant lake.The Devonian transgression began in the South West or South,and spread to the North East or North.Its extent was limited in the Early and early Middle Devonian,but by Givetian time had increased to cover a vast area of South China,including parts of present day(Guangxi,) Hunan,southern Guizhou and eastern Yunnan.As a result the Givetian probably marked the acme of rugose coral development on the South China Plate.Metazoan reefs,with highly diversified reef-building stromatoporoid sponges and both tabulate and rugose corals, flourished.More than 50 such reefs are known from the Givetian of Hunan and Guangxi alone.The rugose coral Endophyllum is well distributed in the Givetian varcus Zone of South China.It commonly occurs together with the brachiopod Stringocephalus,which serves as an index for stratigrapphic correlation(TableⅠ).