中文摘要：

在细胞质、原子的基因之间的 Epistasis 是复杂量的特点的主要基因部件。cytonuclear epistasis 的基因解剖是根本上重要的理解复杂特点的基因建筑学。在这研究，二维的染色体扫描策略被采用评估细胞质的贡献，量的特点 loci (QTL ) ， QTL ? ┤à

英文摘要：

Epistasis between cytoplasmic and nuclear genes is the primary genetic component of complex quantitative traits.Genetic dissection of cytonuclear epistasis is fundamentally important to understand the genetic architecture of complex traits.In this study,a two-dimensional genome scan strategy was employed to evaluate the contribution of cytoplasm,quantitative trait loci （QTL）,QTL×QTL interactions and QTL×QTL×cytoplasm interactions to the phenotypic variation.The p-value and parameter value for each genetic effect were calculated by multiple regression analysis.A stepwise approach was suggested to build confidence in candidate QTL on the basis of q-value estimation,false discovery rate calculation and Bonferroni adjustment.A fine-scale grid scan strategy was proposed for further analysis of peaks of interest.Plant height in maize was used as an example to illustrate the efficiency of the two-dimensional genome scan strategy.